4th instar | 0 | 80 (27) | March-April | Large galls, discolored, blistered leaves |
Pupa | 46 | 310 (66) | April-May | Bud burst, emergence windows and exuviae on leaf undersides |
Adult | 352 | 440 (127) | April-May | Shoot elongation, young leaves with oviposition scars |
Egg | 352 | 748 (104) | May | Same as previous stage |
1st instar | 679 | 1106 (258) | June | Leaf maturation |
2nd instar | 1236 | 2459 (270) | June-September | Gall initiation |
3rd instar | 2443 | 3287 (161) | September-March | Gall enlargement |